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rjin, a Trill candidate to undergo the joining, comes to Deep Space Nine to be Dax's initiate. She has to mentor him and make recommendations if he's worthy to be joined. Arjin seems not happy to be assigned to Dax, knowing what mentor Curzon did to Jadzia. He's trying to do everything according to the normal expectations to please her, but is disappointed to find out she's not an ordinary Trill. On their first away mission in the Gamma Quadrant they accidentally hit a subspace pocket. A strange protoplasmic substance attaches itself to the nacelle. After investigating it on Deep Space Nine, Dax finds out it is a proto-universe. It's expanding and slowly, but surely replacing the present universe. Meanwhile O'Brien and Kira try to battle an infestation of Cardassian Voles. They bite through everything and become more and more a serious threat to the station.

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